腕足动物门


























































腕足動物門
化石时期:下寒武紀–現代

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LingulaanatinaAA.JPG
位於澳大利亞大堡礁斯特德布鲁克岛的鴨嘴海豆芽

科学分类 编辑
界:
动物界 Animalia
亚界:
真后生动物亚界 Eumetazoa
演化支:
副同源异形基因动物 ParaHoxozoa
演化支:
浮浪幼虫样动物 Planulozoa
演化支:
两侧对称动物 Bilateria
演化支:
肾管动物 Nephrozoa
演化支:
原口动物 Protostomia
(未分级):
螺旋動物 Spiralia
总门:
冠輪動物總門 Lophotrochozoa
演化支:
腕足動物 Brachiozoa
门:
腕足動物門 Brachiopoda
Duméril英语André Marie Constant Duméril, 1806[1]

亚门和纲

見內文



多样性[2]

約100個現存屬
約5,000個化石物種。


腕足动物门(學名:Brachiopoda)是动物界的一个门,屬於底栖、有一對硬壳的触手冠海产动物。但與雙殼類動物不同的是:其殼是上、下開合,而不是左、右開合。鉸位在後背部,而前方可開合作捕食或防御。牠們自寒武紀開始演化,现存的种类多分布在高纬度的冷水区。




目录






  • 1 語源


  • 2 特征


  • 3 分類


    • 3.1




  • 4 註釋


  • 5 參考資料


  • 6 延伸閱讀


  • 7 外部連結





語源


腕足動物的學名brachiopod源於古希臘語,由「手臂」(βραχίων)與「足」(πούς)兩部份組成[3]



特征


介壳两枚,大小相等或不等,掩盖背腹两面;介壳的形状和饰纹以及内部器官的构造,是鉴定腕足纲属、种的依据;身体柔软、左右对称;头顶有突出部;上生许多触手,称为“腕足”;消化道呈U字形弯曲,常缺少肛门;具有体腔和后肾。



分類





在西班牙穆爾西亞塞埃欣的早白堊世豪特里維期地層出土的Pygites diphyoides (d'Orbigny, 1849)化石。這件穿孔贝目化石的特色是其殼上位於中央的孔洞。


腕足动物的分類有其獨特的不同之處:現時腕足動物的現存物種與化石物種有着不同的特色。其化石物種的外形變化很大,但其外殼只有少量特徵;而現存物種的外殼變化不大,反而其軟體及外殼均各有特徵。若我們只專注於其中一類特徵,均不能把另一類好好的分類。本門動物也經歷了顯著的趨同進化和逆轉演化(這使較新的組別損失了原來在中間組別出現的特色,又或回復了原來只在較舊的組別出現的特點)。因此,有些分類學的學者認為現時我們仍然未成熟到可以具體把目以上的分類確定,建議只適宜透過自底向上的方法,先確定物種的種屬,然後再把他們分成組[4]。不過,亦有學者不同意這個觀點,認為現時的
然而,其他分類學家相信,物種中某些模式的特點已足夠穩定,值得作更高級別的分類,儘管大家可能對更高級別的分類有不同的看法[4]。以下為現時較多人同意的分類法。





以下為腕足動物門以下的目:




  • 舌形貝型亞門(Linguliformea)


    • 舌形貝綱(Lingulata)

      • †頂孔貝目(Acrotretida,又名乳房贝目)


      • 舌形貝目(Linguilida)

      • †管洞貝目(Siphonotretida)



    • †神父貝綱(Paterinata)
      • †神父贝目(Paterinida)





  • 髑髅贝型亚门(Craniiformea)

    • 頭殻綱(Craniata,又名髑髏貝綱、顱形貝綱)


      • 頭殻目(Craniida,又名髑髏貝目、顱形貝目)

      • †擬髑髏貝目(Craniopsida,又名擬顱形貝目)

      • †三分貝目(Trimerellida)





  • 小嘴貝型亞門(Rhynchonelliformea)


    • 小嘴貝綱(Rhynchonellata)

      • †前正形貝目(Protorthida)

      • †正形贝目(Orthida)

      • †五房贝目(Pentamerida)

      • †无洞贝目(Atrypida,已廢棄的並系群分類)


      • 小嘴贝目(Rhynchonellida)

      • †石燕贝目(Spiriferida)

      • †無窗貝目(Athyridida)

      • †莢采貝目(Spiriferinida)


      • 穿孔贝目(Terebratulida)


      • 鞘貝目(Thecideida)



    • †扭月貝綱(Strophomenata)

      • †扭月贝目(Strophomenida)

      • †长身贝目(Productida)

      • †伯灵贝目(Billingsellida)

      • †直形貝目(Orthotetida)



    • †智利貝綱(Chileata)

      • †智利貝目(Chileida)

      • †目 Dictyonellida



    • †庫圖貝綱(Kutorginata)
      • †庫圖貝目(Kutorginida)


    • †小圓貨貝綱(Obolellata)

      • †小圆货贝目(Obolellida)

      • †目 Naukatida







註釋





  1. ^ Zvyagintsev etc: Brachio fouling (2007).


  2. ^ Cohen: Brachiopoda ELS (2002).


  3. ^ Shorter Oxford English Dictionary (2002),entry "Brachiopod".


  4. ^ 4.04.1 Carlson: Ghosts (2001).




參考資料


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  • Ax, P. Multicellular Animals: Order in Nature - System Made by Man. Multicellular Animals: A New Approach to the Phylogenetic Order in Nature 3. Springer. 2003 [2 November 2009]. ISBN 3-540-00146-8. 


  • Barry, P.L. The Great Dying. Science@NASA. Science and Technology Directorate, Marshall Space Flight Center, NASA. January 28, 2002 [March 26, 2009]. (原始内容存档于2009年4月7日). 


  • Balthasar, U; Butterfield, N.J. Early Cambrian "soft−shelled" brachiopods as possible stem−group phoronids. Acta Palaeontologica Polonica. 2009, 54 (2): 307–314. doi:10.4202/app.2008.0042.  引文使用过时参数coauthors (帮助)


  • Bourlat, S.J; Nielsen, C., Economou, A.D., and Telford, M.J. Testing the new animal phylogeny: A phylum level molecular analysis of the animal kingdom. Molecular Phylogenetics and Evolution. October 2008, 49 (1): 23–31. PMID 18692145. doi:10.1016/j.ympev.2008.07.008.  引文使用过时参数coauthors (帮助)


  • Carlson, S.J. Ghosts of the Past, Present, and Future in Brachiopod Systematics. Journal of Paleontology. November 2001, 75 (6): 1109–1118. doi:10.1666/0022-3360(2001)075<1109:GOTPPA>2.0.CO;2.  引文格式1维护:日期与年 (link)


  • Cohen, B.L. Monophyly of brachiopods and phoronids: reconciliation of molecular evidence with Linnaean classiffication (the subphylum Phoroniformea nov.) (PDF). Proceedings of the Royal Society B: Biological Sciences. 2000, 267 (1440): 225–231 [31 Jan 2010]. doi:10.1098/rspb.2000.0991. 


  • Cohen, B.L; Holmer, L. E. and Lüter, C. The brachiopod fold: a neglected body plan hypothesis (PDF). Palaeontology (Wiley). 2003, 46 (1): 59–65 [2 November 2009]. doi:10.1111/1475-4983.00287.  引文使用过时参数coauthors (帮助)


  • Cohen, B.L. Brachiopoda. Encyclopedia of Life Sciences. John Wiley & Sons, Ltd. 2006. doi:10.1002/9780470015902.a0001614.pub2. 


  • Conway Morris, S; Peel, J. S. Articulated Halkieriids from the Lower Cambrian of North Greenland and their Role in Early Protostome Evolution. Philosophical Transactions of the Royal Society: Biological Sciences. 1995, 347 (1321): 305–358. doi:10.1098/rstb.1995.0029.  引文使用过时参数coauthor (帮助)


  • Cowen, R. Invertebrate paleontology. History of life 3. Wiley-Blackwell. 2000: 408 [2 November 2009]. ISBN 0-632-04444-6. 


  • Doherty, P.J. The Lophophorates. (编) Anderson, D.T. Invertebrate Zoology 2. Oxford University Press. 2001: 356–363. ISBN 0-19-551368-1. 


  • Dunn, C.W; Hejnol, A.; Matus, D.Q.; Pang, K.; Browne, W.E.; Smith, S.A.; Seaver, E.; Rouse, G.W.; Obst, M. Broad phylogenomic sampling improves resolution of the animal tree of life. Nature. 2008, 452 (7188): 745–749 [3 Feb 2010]. Bibcode:2008Natur.452..745D. PMID 18322464. doi:10.1038/nature06614. 


  • Emig, C.C. Ecology of Inarticulated Brachiopods. (编) Kaesler, R.L. [[Treatise on Invertebrate Paleontology]] (PDF) Part H. Geological Society of America. 1997: 473–481 [14 November 2009].  网址-维基内链冲突 (帮助)


  • Fortey, R.A. How to recognize fossils. Fossils: the key to the past. Harvard University Press. 1991: 61–63 [16 November 2009]. ISBN 0-674-31135-3. 


  • Gaines, R.R; Droser. Animal proxies - Invertebrates. (编) Gornitz, V. Encyclopedia of paleoclimatology and ancient environments. M.L. Springer. 2009: 11 [15 November 2009]. ISBN 1-4020-4551-4.  引文使用过时参数coauthors (帮助)


  • Giribet, G; Distel, D.L., Polz, M., Sterrer, W., and Wheeler W. Triploblastic relationships with emphasis on the acoelomates and the position of Gnathostomulida, Cycliophora, Plathelminthes, and Chaetognatha: a combined approach of 18S rDNA sequences and morphology. Systemic Biology. Sep 2000, 49 (3): 539–62.  引文使用过时参数coauthors (帮助)


  • Gould, S.J; Calloway, C.B. Clams and Brachiopods - Ships that Pass in the Night. Paleobiology (Paleontological Society). 1980, 6 (4): 383–396. JSTOR 2400538.  引文使用过时参数coauthors (帮助)


  • Halanych, K.M. The new view of animal phylogeny (PDF). Annual Review of Ecology, Evolution, and Systematics. 2004, 35: 229–256 [21009-04-17]. doi:10.1146/annurev.ecolsys.35.112202.130124.  请检查|access-date=中的日期值 (帮助)


  • Helmkampf, M; Bruchhaus, I., and Hausdorf, B. Phylogenomic analyses of lophophorates (brachiopods, phoronids and bryozoans) confirm the Lophotrochozoa concept. Proceedings of the Royal Society B: Biological Sciences. August 2008, 275 (1645): 1927–1933. PMC 2593926. PMID 18495619. doi:10.1098/rspb.2008.0372.  引文使用过时参数coauthors (帮助)


  • Iyengar, E.V. Kleptoparasitic interactions throughout the animal kingdom and a re-evaluation, based on participant mobility, of the conditions promoting the evolution of kleptoparasitism. Biological Journal of the Linnean Society (Wiley InterScience). March 2008, 93 (4): 745–762. doi:10.1111/j.1095-8312.2008.00954.x. 


  • Knoll, A.H., Bambach, R.K., Payne, J.L., Pruss, S., and Fischer, W.W. Paleophysiology and end-Permian mass extinction (PDF). Earth and Planetary Science Letters. 2007, 256 (3–4): 295–313 [2008-07-04]. Bibcode:2007E&PSL.256..295K. doi:10.1016/j.epsl.2007.02.018. 


  • Kowalewski, M; Hoffmeister, A.P., Baumiller, T.K., and Bambach, R.K. Secondary Evolutionary Escalation Between Brachiopods and Enemies of Other Prey. Science. June 2005, 308 (5729): 1774–1777. doi:10.1126/science.1113408.  引文使用过时参数coauthors (帮助)


  • ITIS: Brachiopoda. Integrated Taxonomic Information System. [16 Nov 2009]. 


  • Milsom, C; Rigby, S. Brachiopods. Fossils at a Glance. John Wiley and Sons. 2009: 37 [16 November 2009]. ISBN 1-4051-9336-0.  引文使用过时参数coauthors (帮助)


  • Nielsen, C. The Phylogenetic Position of Entoprocta, Ectoprocta, Phoronida, and Brachiopoda. Integrative and Comparative Biology. 2002, 42 (3): 685–691. PMID 21708765. doi:10.1093/icb/42.3.685. 


  • Nielsen, C. Larval and adult brains. Evolution & Development (Wiley InterScience). 2005, 7 (5): 483–489. doi:10.1111/j.1525-142X.2005.05051.x. 


  • Parkinson, D; Curry, G.B., Cusack, M., and Fallick, A.E. Shell structure, patterns and trends of oxygen and carbon stable isotopes in modern brachiopod shells. Chemical Geology (Elsevier). 2005, 219 (1–4): 193–235. doi:10.1016/j.chemgeo.2005.02.002.  引文使用过时参数coauthors (帮助)


  • Peterson, K.J; Eernisse, D.J. Animal phylogeny and the ancestry of bilaterians: inferences from morphology and 18S rDNA gene sequences. Evolution and Development. May 2001, 3 (3): 170–205. PMID 11440251. doi:10.1046/j.1525-142x.2001.003003170.x.  引文使用过时参数coauthors (帮助)


  • Riisgård, H.U; Nielsen, C., and Larsen, P.S. Downstream collecting in ciliary suspension feeders: the catch-up principle (PDF). Marine Ecology Progress Series. 2000, 207: 33–51 [12 September 2009]. doi:10.3354/meps207033.  引文使用过时参数coauthors (帮助)


  • de Rosa, R. Molecular Data Indicate the Protostome Affinity of Brachiopods (PDF). Systematic Biology. 2001, 50 (6): 848–859 [27 Jan 2010]. PMID 12116636. doi:10.1080/106351501753462830. 


  • Ruppert, E.E; Fox, R.S., and Barnes, R.D. Invertebrate Zoology 7. Brooks / Cole. 2004. ISBN 0-03-025982-7.  引文使用过时参数coauthors (帮助)


  • Christian B. Skovsted, Glenn A. Brock, John R. Paterson, Lars E. Holmer, Graham E. Budd. The scleritome of Eccentrotheca from the Lower Cambrian of South Australia: Lophophorate affinities and implications for tommotiid phylogeny. Geology. 2008-02-01, 36 (2) [2018-04-03]. ISSN 0091-7613. doi:10.1130/g24385a.1 (英语). 


  • Trumble, W; Brown, L. Shorter Oxford English Dictionary. Oxford University Press, USA. 2002. ISBN 0-19-860457-2.  引文使用过时参数coauthors (帮助)


  • Introduction to the Lingulata. University of California Museum of Paleontology. [16 Nov 2009]. 


  • Fossil Range Chart of Brachiopods. University of California Museum of Paleontology. [16 Nov 2009]. 


  • Introduction to the Deuterostomia. University of California Museum of Paleontology. [27 January 2010]. 


  • Ushatinskaya, G.T. Origin and dispersal of the earliest brachiopods. Paleontological Journal (Springer Verlag). 2008, 42 (8): 776–791. doi:10.1134/S0031030108080029. 


  • Valentine, J.W. Cleavage patterns and the topology of the metazoan tree of life. Proceedings of the National Academy of Sciences. 1997, 94 (15): 8001–8005. Bibcode:1997PNAS...94.8001V. PMC 21545. PMID 9223303. doi:10.1073/pnas.94.15.8001. 


  • Vermeij, G.T. Inequality and the Directionality of History. The American Naturalist (The American Society of Naturalists). March 1999, 153 (3): 243–253. JSTOR 2463821. 


  • Williams, A; Carlson, C.H.C; Brunton, S.J. Outline of Suprafamilial Classification and Authorship. (编) Williams, A., Carlson, C.H.C, and Brunton, S.J. Brachiopoda. Treatise on Invertebrate Paleontology (Geological Society of America and The University of Kansas). 2000: xxxix–xlv. ISBN 0-8137-3108-9. 


  • Wood, T.S; Lore. The higher phylogeny of phylactolaemate bryozoans inferred from 18S ribosomal DNA sequences. (编) Moyano, H. I., Cancino, J. M. and Wyse-Jackson, P.N. Bryozoan Studies 2004: Proceedings of the 13th International Bryozoology Association. M. London: Taylor & Francis Group. 2005: 361–367.  引文使用过时参数coauthors (帮助)


  • Zvyagintsev, A.Y; Radashevsky, V. I., and Kashin, I. A. First record of a brachiopod (Brachiopoda: Terebrataliidae) in the fouling of hydrotechnical installations in Peter the Great Bay, Sea of Japan. Russian Journal of Marine Biology. August 2007, 33 (4): 264–266. doi:10.1134/S1063074007040098.  引文使用过时参数coauthors (帮助)




延伸閱讀



  • Moore, R.C.; Lalicker, C.G., and Fischer, A.G. Invertebrate Fossils. Mcgraw-Hill College. June 1952. ISBN 0-07-043020-9.  引文使用过时参数coauthors (帮助)


外部連結







  • UC-Berkeley Museum of Paleontology

  • Palaeos Brachiopoda

  • BrachNet

  • Information from the Kansas Geological Survey

  • site of R.Filippi

  • Brachiopoda World Database





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